Swimming
Indicator of swimming activity and energetic demand
Increases linearly with swimming speed through the sustained aerobic range.
Strong correlation between tail beat frequency and metabolic rate.
At high speeds, both tail beat frequency and amplitude increase to increase thrust.

Current speed ▲ Increases frequency
Tail beat frequency increases when fish swim against faster currents.

Temperature ▲ Increases frequency
Maximum achievable tail beat frequency is higher in warmer temperatures. 
Fed status ◆ No effect
Fed and fasted fish show the same tail beat frequency across current speeds, but fasted fish have lower metabolic rates throughout.
